2001 Directory                                                        Faculty Information Page 2001
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(Photo of Dr. Hulse)
(IUP Seal)

Dr. Arthur C. Hulse

Professor of Biology

Indiana Univ. of PA


Electronic Mail Address



Office: (724) 357-2279
Biology Department Office: (724) 357-2352
FAX: (724) 357-5700


Semester Schedule

Office Hours

Teaching Schedule


IUP Campus Address

314 Weyandt Hall
Department of Biology
Indiana University of Pennsylvania
Indiana, Pennsylvania 15705

Professional Interests and Formal Education
[ Publications ]

Field: Systematic and ecological herpetology.

         Much of my current research is directed towards the long-term population biology and reproductive ecology of aquatic salamanders.

         I am investigating the population structure, activity, and reproduction of a metapopulation of the red-spotted newt (
Notophthalmus viridescens) in a series of abandoned beaver ponds. I have also instituted a research program into the ecology and conservation status of the eastern hellbender (Cryptobranchus alleganiensis alleganiensis ). This involves a long-term mark-recapture study of populations of hellbenders in three streams with different histories of environmental impact as well as a state wide sampling of hellbenders to determine their overall distribution within Pennsylvania.

         In addition I am also working on the reproductive biology of the redbelly snake (
Storeria occipitomaculata ) and the ringneck snake (Diadophis punctatus ). These studies are designed to examine variation in reproductive parameters within a species due to temporal and geographic factors.

         I am also writing a book dealing with the natural history of the amphibians and reptiles of Pennsylvania in conjunction with the Carnegie Museum of Natural History.

B.S.   - 1967 Bloomfield College
M.S. - 1970 Arizona State University
Ph.D. - 1974 Arizona State University

Biology Department Web Site Maintainer: Dr. Ray L. Winstead
Direct e-mail Link: RWinstea@grove.iup.edu